Marine Environmental Biology, Department of Biological Sciences, University of Southern California, 3616 Trousdale Pkwy., Los Angeles, California, 90089, USA

now at: Northeast Fisheries Science Center/NOAA, 212 Rogers Ave., Milford, Connecticut, 06460, USA

Marine Environmental Biology, Department of Biological Sciences, University of Southern California, 3616 Trousdale Pkwy., Los Angeles, California, 90089, USA

NIWA Centre for Chemical and Physical Oceanography, Department of Chemistry, University of Otago, P.O. Box 56, Dunedin, New Zealand

National Institute of Water and Atmosphere, 301 Evans Bay Parade, Greta Point, Wellington, New Zealand

Bigelow Laboratory for Ocean Sciences, 180 McKown Point Road., P.O. Box 475, West Boothbay Harbor, Maine, 04575–0475, USA

Dept. of Microbiology, The University of Tennessee, Knoxville, Tennessee, 37996, USA

Marine Environmental Biology, Department of Biological Sciences, University of Southern California, 3616 Trousdale Pkwy., Los Angeles, California, 90089, USA

Research School of Earth Sciences, Australian National University, ACT 0200, Australia

Marine Environmental Biology, Department of Biological Sciences, University of Southern California, 3616 Trousdale Pkwy., Los Angeles, California, 90089, USA